Stone Pine

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Stone Pine
Pinus pinea
Stone Pine forest
Costa de la Luz, Spain
Conservation status
Scientific classification
Kingdom:Plantae
Division:Pinophyta
Class:Pinopsida
Order:Pinales
Family:Pinaceae
Genus:Pinus
Subgenus:Pinus
Species:P. pinea
Binomial name
Pinus pinea
L.
 
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Stone Pine
Pinus pinea
Stone Pine forest
Costa de la Luz, Spain
Conservation status
Scientific classification
Kingdom:Plantae
Division:Pinophyta
Class:Pinopsida
Order:Pinales
Family:Pinaceae
Genus:Pinus
Subgenus:Pinus
Species:P. pinea
Binomial name
Pinus pinea
L.

The Stone Pine (Pinus pinea) is also called Italian Stone Pine, Umbrella Pine (not to be confused with Japanese Umbrella pine, Sciadopitys verticillata), and Parasol Pine. It is in the pine family Pinaceae and occasionally listed under the invalid name Pinus sativa. The tree is native to the Mediterranean region. It occurs in Southern Europe, North Africa, and the Levant.

Stone pines have been used and cultivated for their edible pine nuts since prehistoric times. They are widespread in horticultural cultivation as ornamental trees, planted in gardens and parks around the world. This plant has gained the Royal Horticultural Society's Award of Garden Merit.[1]

Contents

Distribution

The prehistoric range of Pinus pinea included North Africa in the Sahara Desert and Maghreb regions during a more humid climate period, in present day Morocco, Algeria, Tunisia, and Libya. Its contemporary natural range is in the Mediterranean forests, woodlands, and scrub biome ecoregions and countries, including:

Southern Europe

The Iberian conifer forests ecoregion of the Iberian Peninsula in Spain and Portugal; the Italian sclerophyllous and semideciduous forests ecoregion in France and Italy; the Tyrrhenian-Adriatic sclerophyllous and mixed forests ecoregion of southern Italy, Sicily, and Sardinia; the Illyrian deciduous forests of the eastern coast of the Ionian and Adriatic Seas in Croatia and Albania; and the Aegean and Western Turkey sclerophyllous and mixed forests ecoregion of the southern Balkan Peninsula in Greece.

In Greece, although rare[2], an extensive Stone Pine forest exists in western Peloponnese at Strofylia[3] on the peninsula separating the Kalogria Lagoon from the Mediterranean Sea. This coastal forest is at least 8 miles long, with dense and tall stands of Pinus pinea mixed with Pinus halepensis.[4] Currently, Pinus halepensis is outcompeting Stone Pines in many locations of the forest.[5] Another location in Greece is at Koukounaries on the northern Aegean island of Skiathos at the southwest corner of the island. This is a half a mile long dense stand of Stone and Aleppo Pines that lies between a lagoon and the Aegean Sea.[6] A fine-textured sand beach lies between the Koukounaries forest and the sea.

Western Asia

In Western Asia, the Eastern Mediterranean conifer-sclerophyllous-broadleaf forests ecoregion in Turkey; and the Southern Anatolian montane conifer and deciduous forests ecoregion in Lebanon, Syria, and northern Israel

northern Africa

The Mediterranean woodlands and forests ecoregion of North Africa, in Morocco and Algeria

Description

Pinus pinea: juvenile (left) — adult foliage (right)
Pinus pinea: the bark's vertical texture in close-up
Pinus pinea: the cone and seeds

The Stone pine is a coniferous evergreen tree growing to 12–20 metres (39–66 ft) in height, and can exceed 25 metres (82 ft) height. In youth, it is a bushy globe, in mid-age an umbrella canopy on a thick trunk, and, in maturity, a broad and flat crown 40–60 metres (130–200 ft) in width. The bark is thick, red-brown and deeply fissured into broad vertical plates.

Foliage

The flexible mid-green leaves are needle-like, in bundles of two, and are 10–20 centimetres (3.9–7.9 in) long (exceptionally up to 30 centimetres (12 in)). Young trees up to 5–10 years old bear juvenile leaves, which are very different, single (not paired), 2–4 centimetres (0.79–1.6 in) long, glaucous blue-green; the adult leaves appear mixed with juvenile leaves from the fourth or fifth year on, replacing it fully by around the tenth year. Juvenile leaves are also produced in regrowth following injury, such as a broken shoot, on older trees.

Cones

The cones are broad, ovoid, 8–15 centimetres (3.1–5.9 in) long, and take 36 months to mature, longer than any other pine. The seeds (pine nuts, piñones, pinhões, pinoli, or pignons) are large, 2 centimetres (0.79 in) long, and pale brown with a powdery black coating that rubs off easily, and have a rudimentary 4–8 millimetres (0.16–0.31 in) wing that falls off very easily. The wing is ineffective for wind dispersal, and the seeds are animal-dispersed, originally mainly by the azure-winged magpie, but in recent history, very largely by humans.

Cultivation

Food

Pinus pinea has been cultivated extensively for at least 6,000 years for the edible pine nuts. These have been trade items since early historic times. It has been cultivated throughout the Mediterranean region for so long that it has naturalized, and is often considered native beyond its natural range.

Ornamental tree

In Italy, the Stone Pine has been an aesthetic landscape element since the Italian Renaissance garden period. In the 1700s, P. pinea began being introduced as an ornamental tree to other Mediterranean climate regions of the world, and is now often found in gardens and parks in South Africa, California, and Australia. It has naturalized beyond cities in South Africa to the extent that it is listed as an invasive species there. It is also planted in western Europe up to southern Scotland, and on the East Coast of the United States up to New Jersey.

Small specimens are used for Bonsai, and also grown in large pots and planters. The year-old seedlings are seasonally available as 20–30 centimetres (7.9–12 in) tall table-top Christmas trees.

Mid-age tree with umbrella canopy, in Catalonia, Spain.
Mature tree with flatter crown, in Tuscany, Italy

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Pests

The introduced western conifer seed bug (Leptoglossus occidentalis) was accidentally imported with timber to northern Italy in the late 1990s from western USA, and has spread across Europe as an invasive pest species since then. It feeds on the sap of developing conifer cones throughout its life, and its sap-sucking causes the developing seeds to wither and misdevelop. It has destroyed most of the pine nut seeds in Italy, threatening P. pinea in its native habitats there.[7]

See also

References