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|Seeds from various perennial flowers.|
A perennial plant or simply perennial (from Latin per, meaning "through", and annus, meaning "year") is a plant that lives for more than two years. The term is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widely used to distinguish plants with little or no woody growth from trees and shrubs, which are also technically perennials.
Perennials, especially small flowering plants, that grow and bloom over the spring and summer, die back every autumn and winter, and then return in the spring from their root-stock, are known as herbaceous perennials. However, depending on the rigors of local climate, a plant that is a perennial in its native habitat, or in a milder garden, may be treated by a gardener as an annual and planted out every year, from seed, from cuttings or from divisions. Tomato vines, for example, live several years in their natural tropical/subtropical habitat but are grown as annuals in temperate regions because they don't survive the winter.
There is also a class of evergreen, or non-herbaceous, perennials, including plants like Bergenia which retain a mantle of leaves throughout the year. An intermediate class of plants is known as subshrubs, which retain a vestigial woody structure in winter, e.g. Penstemon. The local climate may dictate whether plants are treated as shrubs or perennials. For instance, in colder temperate climates, many shrubby varieties of Fuchsia are cut to the ground to protect them from winter frosts.
Perennial plants can be short-lived (only a few years) or they can be long-lived, as are some woody plants like trees. They include a wide assortment of plant groups from ferns and liverworts to the highly diverse flowering plants like orchids and grasses.
Perennials typically grow structures that allow them to adapt to living from one year to the next through a form of vegetative reproduction rather than seeding. These structures include bulbs, tubers, woody crowns, rhizomes plus others. They might have specialized stems or crowns that allow them to survive periods of dormancy over cold or dry seasons during the year. Annuals produce seeds to continue the species as a new generation while the growing season is suitable, and the seeds survive over the cold or dry period to begin growth when the conditions are again suitable.
Many perennials have developed specialized features that allow them to survive extreme climatic and environmental conditions. Some have adapted to survive hot and dry conditions or cold temperatures. Those plants tend to invest a lot of resource into their adaptations and often do not flower and set seed until after a few years of growth. Many perennials produce relatively large seeds, which can have an advantage, with larger seedlings produced after germination that can better compete with other plants. Some annuals produce many more seeds per plant in one season, while some (polycarpic) perennials are not under the same pressure to produce large numbers of seeds but can produce seeds over many years.
In warmer and more favorable climates, perennials grow continuously. In seasonal climates, their growth is limited to the growing season.
In some species, perennials retain their foliage all year round; these are evergreen perennials. Other plants are deciduous perennials, for example, in temperate regions a perennial plant may grow and bloom during the warm part of the year, with the foliage dying back in the winter. In many parts of the world, seasonality is expressed as wet and dry periods rather than warm and cold periods, and deciduous perennials lose their leaves in the dry season.
With their roots protected below ground in the soil layer, perennial plants are notably tolerant of wildfire. Herbaceous perennials are also able to tolerate the extremes of cold in temperate and Arctic winters, with less sensitivity than trees or shrubs.
Although most of humanity is fed by seeds from annual grain crops, perennial crops provide numerous benefits. Perennial plants often have deep, extensive root systems which can hold soil to prevent erosion, capture dissolved nitrogen before it can contaminate ground and surface water, and outcompete weeds (reducing the need for herbicides). These potential benefits of perennials have resulted in new attempts to increase the seed yield of perennial species, which could result in the creation of new perennial grain crops. Some examples of new perennial crops being developed are perennial rice and intermediate wheatgrass. The Land Institute estimates that profitable, productive perennial grain crops will take at least 25 years to achieve.
|Seeds from various perennial flowers.|
Perennial plants dominate many natural ecosystems on land and in fresh water, with only a very few (e.g. Zostera) occurring in shallow sea water. Herbaceous perennial plants are particularly dominant in conditions too fire-prone for trees and shrubs, e.g., most plants on prairies and steppes are perennials; they are also dominant on tundra too cold for tree growth. Nearly all forest plants are perennials, including the trees and shrubs.
Perennial plants are usually better competitors than annual plants, especially under stable, resource-poor conditions. This is due to the development of larger root systems which can access water and soil nutrients deeper in the soil and to earlier emergence in the spring.
The following perennial plants are used as herbs:
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Many vegetable plants can grow as perennials in tropical climates, but die in cold weather. Some of the more completely perennial vegetables are: