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Fire ant is the common name for several species of ants in the genus Solenopsis. They are however only a minority in the genus, which includes over 286 species of Solenopsis worldwide. Solenopsis are stinging ants and most of their common names reflect this, for example, ginger ants and tropical fire ants. Many species also are called red ants because of their light brown colour, though species of ants in many other genera are similarly named for similar reasons. Examples include Myrmica rubra and Pogonomyrmex barbatus.
None of these names applies in all countries nor to all species of Solenopsis, nor only to Solenopsis species; for example the colloquial names for several species of weaver ants in the genus Oecophylla in Southeast Asia include "fire ants" because of their red color and painful sting; the two genera however are not closely related.
The bodies of mature fire ants, like the bodies of all typical mature insects, are divided into three sections: the head, the thorax, and the abdomen, with three pairs of legs and a pair of antennae. Fire ants of those species invasive in the United States can be distinguished from other ants locally present, by their copper brown head and body with a darker abdomen. The worker ants are blackish to reddish, and their size varies from 2 mm to 6 mm (0.12 in to 0.24 in). In an established nest these different sizes of ants all are present at the same time.
Many ants bite, and formicine ants can cause irritation by spraying formic acid; myrmecine ants like fire ants have a dedicated venom-injecting sting, which injects an alkaloid venom, as well as mandibles for biting.
A typical fire ant colony produces large mounds in open areas, and feeds mostly on young plants, seeds and sometimes crickets. Fire ants often attack small animals and can kill them. Unlike many other ants, which bite and then spray acid on the wound, fire ants bite only to get a grip and then sting (from the abdomen) and inject a toxic alkaloid venom called Solenopsin, a compound from the class of piperidines. For humans, this is a painful sting, a sensation similar to what one feels when burned by fire (hence the name) and the after effects of the sting can be deadly to sensitive people. The liquid is both insecticidal and antibiotic.
Fire ants are more aggressive than most native species and so have pushed many species away from their local habitat. These ants are renowned for their ability to survive extreme conditions. They do not hibernate, but can survive cold conditions as low as -9 °C (16° F) despite originating from South America.
Fire ants nest in the soil, often near moist areas, such as river banks, pond shores, watered lawns and highway shoulder. Usually, the nest will not be visible, as it will be built under objects such as timber, logs, rocks, or bricks. If there is no cover for nesting, dome-shaped mounds will be constructed, but these are usually only found in open spaces, such as fields, parks and lawns. These mounds can reach heights of 40 cm (15.7 in), and can also be as deep as 1.5 meters (5 feet).
Colonies are founded by small groups of queens or single queens. Even if only one queen survives, within a month or so, the colony can expand to thousands of individuals. Some colonies may be polygynous (having multiple queens per nest).
Fire ant queens, the reproductive females in their colony, also are generally the largest. Their primary function is reproduction; fire ant queens may live as many as 6 to 7 years and can produce up to 3,500 eggs per day. That adds up to roughly 9 million eggs produced during a single queen's lifetime. Young, virgin fire ant queens have wings (as do male fire ants), but rip them off after mating.
Males mate with the queen. They die immediately after mating.
There are other types of roles in an ant colony like the workers and the soldier ants. The soldier ants are known for their larger and more powerful mandibles while the worker takes care of regular tasks (the main tasks in a colony are caring for the eggs/larvae/pupae, cleaning the nest, and foraging for food).
Although most fire ant species do not bother people and are not invasive, Solenopsis invicta, known in the United States as the red imported fire ant (or RIFA) is an invasive pest in many areas of the world, notably the United States, Australia, the Philippines, China and Taiwan. The RIFA was believed to have been accidentally introduced to these countries via shipping crates.
In the US the FDA estimates that more than US$5 billion is spent annually on medical treatment, damage, and control in RIFA-infested areas. Furthermore, the ants cause approximately $750 million in damage annually to agricultural assets, including veterinarian bills and livestock loss, as well as crop loss. Over 40 million people live in RIFA-infested areas in the southeastern United States. Between 30 and 60% of people living in fire ant-infested areas are stung each year. Since September 2004 Taiwan has been seriously affected by the red fire ant. The US, Taiwan and Australia all have ongoing national programs to control or eradicate the species, but, except for Australia, none have been especially effective. In Australia, an intensive program costing A$175 million had reportedly by February 2007 eradicated 99% of fire ants from the sole infestation occurring in south-east Queensland, however by July 2013 multiple sites west of Brisbane were confirmed, including the Lockyer Valley, Muirlea and Goodna
According to a study published in 2009, it only took seventy years for the lizards in parts of the United States to adapt to the ant's presence – they now have longer legs and new behaviors that aid them in escaping from the danger.
The venom of fire ants is composed of alkaloids derived from piperidine (see Solenopsis saevissima). Some people are allergic to the venom, and as with many allergies, may experience anaphylaxis, which requires emergency treatment. Other problems are concentrated to the site of the sting.
The sting swells into a bump, which can cause much pain and irritation, especially when several stings are in the same place. The bump often forms into a white pustule, which can become infected if scratched, but if left alone will usually flatten within a few days. The pustules are obtrusive and uncomfortable while active and, if they become infected, can cause scarring.
First aid for fire ant stings includes external treatments and oral medicines. There are also many home remedies of varying efficacy, including immediate application of urine or aloe vera gel, the latter of which is also often included in over-the-counter creams that also include medically tested and verified treatments.
External, topical treatments include the anesthetic benzocaine, the antihistamine diphenhydramine, and the corticosteroid hydrocortisone. Antihistamines or topical corticosteroids may help reduce the itching.
Oral medicine include antihistamines.
Severe allergic reactions to fire ant stings, including severe chest pain, nausea, severe sweating, loss of breath, serious swelling, and slurred speech, can be fatal if not treated.
Phorid flies, or Phoridae, are a large family of small, hump-backed flies somewhat smaller than vinegar flies; two species in this family (Pseudacteon tricuspis and Pseudacteon curvatus) are parasitoids of the red imported fire ant in its native range in South America. Some 110 species of the genus Pseudacteon, or ant-decapitating flies, have been described. Members of Pseudacteon reproduce by laying eggs in the thorax of the ant. The first instar larvae migrates to the head, then develops by feeding on the hemolymph, muscle tissue, and nervous tissue. After about two weeks, they cause the ant's head to fall off by releasing an enzyme that dissolves the membrane attaching the ant's head to its body. The fly pupates in the detached head capsule, emerging two weeks later.
Pseudacteon flies appear to be important ecological constraints on Solenopsis species and they have been introduced throughout the southern United States, starting with Travis, Brazos, and Dallas counties in Texas, as well as Mobile, Alabama, where the ants first entered North America.
The Venus flytrap, a carnivorous plant, is native to North and South Carolina in the United States. About 33% of the prey of the Venus Flytrap are ants of various species. They lure their prey with a sweet sap. Once the prey has entered the trap and within about 30 seconds has touched two or three "trigger hairs", bristles on the surface of the trap, the leaf closes around the prey, confining it behind the "teeth" on its perimeter, and digests it. Other carnivorous plants, such as Sundews (Drosera) and various kinds of pitcher plants also trap many ants, but in contrast to the effectiveness of relevant species of Phorid flies, there is no evidence that predation by carnivorous plants is in practice a significant drain on populations of Solenopsis species.
A checklist of species of Solenopsis is accessible at Antwiki 
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