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Toes are the digits of the foot of a tetrapod. Animal species such as cats that walk on their toes are described as being digitigrade. Humans, and other animals that walk on the soles of their feet, are described as being plantigrade; unguligrade animals are those that walk on hooves at the tips of their toes.
The toe refers to a part of the human foot, with five toes present on each human foot. Each toe consists of three phalanx bones, the proximal, middle, and distal, with the exception of the big toe (Latin: Hallux). The hallux only contains two phalanx bones, the proximal and distal. The phalanx bones of the toe join to the metatarsal bones of the foot at the interphalangeal joints.:557 Outside the hallux bones is skin, and present on all five toes is a toenail.
The toes are, from medial to lateral:
With the exception of the hallux, toe movement is generally governed by action of the flexor digitorum brevis and extensor digitorum brevis muscles. These attach to the sides of the bones,:572–5 making it impossible to move individual toes independently. Muscles between the toes on their top and bottom also help to bring abduct and adduct the toes.:579 The hallux and little toe have unique muscles:
Sensation to the skin of the toes is provided by five nerves. The superficial fibular nerve supplies sensation to the top of the toes, except between the hallux and second toe, which is supplied by the deep fibular nerve, and the outer surface of the fifth toe, supplied by the sural nerve. Sensation to the bottom of the toes is supplied by the medial plantar nerve, which supplies sensation to the great toe and inner three-and-a-half toes, and the lateral plantar nerve, which supplies sensation to the little toe and half of the sensation of the fourth toe.:583
In humans, the hallux is usually longer than the second toe. There is an inherited trait in humans, where the dominant gene causes a longer second toe ("Morton's toe" or "Greek foot") while the homozygous recessive genotype presents with the more common trait: a longer hallux. People with the rare genetic disease fibrodysplasia ossificans progressiva characteristically have a short hallux which appears to turn inward, or medially, in relation to the foot.
The hallux may not be the longest toe on the foot of some individuals. The name stems from the Latin for the same, similar to the pollex, which refers to the thumb, the corresponding digit of the hand. The fifth toe is the outermost usually the smallest toe of the foot.
Forefoot shape, including toe shape, exhibits significant variation among people; these differences can be measured and have been statistically correlated with ethnicity. Such deviations may affect comfort and fit for various shoe types. Research conducted by Freedman for the U.S. Army indicated that larger feet may still have smaller arches, toe length, and toe-breadth.
The human foot consists of multiple bones and soft tissues which support the weight of the upright human. Specifically, the toes assist the human while walking, providing balance, weight-bearing, and thrust during the gait.
Long-term use of improperly sized shoes can cause misalignment of toes, as well as other orthopedic problems.
Morton's neuroma commonly results in pain and numbness in the third and fourth toes of the sufferer, due to it affecting the nerve between the third and fourth metatarsal bones. If this occurs, the symptoms will be felt up left-hand side of the third toe (and the right-hand side of the fourth toe.
Deformities of the foot include hammer toe, trigger toe, and claw toe. Hammer toe can be described as an abnormal contraction or “buckling” of a toe. This is done by a partial or complete dislocation of one of the joints, which form the toes. Since the toes are deformed further, these may press against a shoe and cause pain. Deformities of the foot can also be caused by rheumatoid arthritis and diabetes mellitus. Deformities may predispose to ulcers and pain, especially when shoe-wearing.
A common problem involving the big toe is the formation of bunions. These are structural deformities of the bones and the joint between the foot and big toe, and may be painful. Similar deformity involving the fifth toe is described as tailor's bunion or bunionette.
A favourable option for the reconstruction of missing adjacent fingers/multiple digit amputations, i.e. such as a metacarpal hand reconstruction, is to have a combined second and third toe transplantation. Third and fourth toe transplantation are also good.
In classical Latin hallex, allex, hallus  and allus, with genitive (h)allicis and (h)alli, are used to refer to the big toe. The form hallux (genitive, hallucis) currently in use is however a blend word of the aforementioned forms.
While the thumb is often mentioned as one of the signature characteristics in humans, this manual digit remains partially primitive and is actually present in all primates. In humans, the most derived digital feature is the hallux.
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