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The dun gene is a dilution gene that affects both red and black pigments in the coat color of a horse. The dun gene has the ability to affect the appearance of all black, bay, or chestnut-based horses to some degree by lightening the base body coat and suppressing the underlying base color to the mane, tail, legs, and "primitive markings".
The classic dun is a gray-gold or tan, characterized by a body color ranging from sandy yellow to reddish-brown. A dun horse always has a dark stripe down the middle of its back, a tail and mane darker than the body coat, and usually darker face and legs. Other duns may appear a light yellowish shade, or a steel gray, depending on the underlying coat color genetics. Manes, tails, primitive markings, and other dark areas are usually the shade of the undiluted base coat color.
The dun allele is a simple dominant, so the phenotype of a horse with either one copy or two copies of the gene is dun. It has a stronger effect than other dilution genes, such as the silver dapple gene, which acts only on black-based coats, or the cream gene, an incomplete dominant which must be homozygous to be fully expressed, and when heterozygous is only visible on bay and chestnut coats, and then to a lesser degree.
The dun gene also is characterized by primitive markings, which are darker than the body color. Primitive markings include:
Dorsal striping does not guarantee the horse carries the dun gene. A countershading gene can also produce faint dorsal striping, even in breeds such as the Arabian horse or the Thoroughbred, where the dun gene is not known to be carried in the gene pool. A primary characteristic of the dun gene is the dorsal stripe, and most duns also have visual leg striping. The shoulder stripes are less common and often fainter, but usually visible on horses with a short summer coat.
The dun coat color is thought to be a primitive trait in the horse. This is because equines appearing in prehistoric cave paintings are dun, and because several closely related species in the genus Equus are known to have been dun. These species include both subspecies of Equus ferus (the extinct tarpan and the extant, but endangered Przewalski's horse), the extinct Equus lambei, and the extant onager and kiang.
The dun gene has a stronger dilution effect on the body than the mane, tail, legs, and primitive markings, so lightens the body coat more. This explains why points on a dun are a shade darker than the coat, or in the case of a "classic" dun, the mane, tail, and legs often are black or only slightly diluted.
Since the dun gene, when on a "bay dun" horse, can closely resemble buckskin, in that both colors feature a light-colored coat with a dark mane and tail, classic duns frequently are confused with buckskins. The difference between these two colors is the dun as a tan color, somewhat duller than the more cream or gold buckskin, and duns also possess primitive markings. Some buckskins do show countershading, but it is not related to the primitive markings of dun factor horses.
Genetically, a bay dun is a bay horse with the dun gene that causes the lighter coat color and the primitive markings. A buckskin is bay horse with the addition of the cream gene, causing the coat color to be diluted from red to gold, often without primitive markings.
A red dun may also be confused with a perlino , which is genetically a bay horse with two copies of the cream gene, which creates a horse with a cream-colored body but a reddish mane and tail. However, perlinos usually are significantly lighter than a red dun and generally have blue eyes.
To further confuse matters, it is possible for a horse to carry both dun and cream dilution genes; such horses with golden buckskin coloring and a complete set of primitive markings are referred to as a "buckskin dun" or a "dunskin". In the Fjord horse, duns that also carry the creme dilution are called uls dun or white dun (ulsblakk) and yellow dun (gulblakk) by their respective coat colors. On such horses, the light-shaded primitive markings are most noticeable during the summer months when the winter hair sheds.
Countershading is a usually a darker shade of the body color rather than the near-black of primitive markings on bay duns, but it may be harder to differentiate between countershading and a dorsal stripe on light-colored horses such as red duns. In such cases, pedigree analysis, DNA testing, studying possible offspring, and the presence of other primitive markings are used to determine whether a horse is a dun.
The three primary dun varieties usually occur in proportion to the occurrence of the corresponding base colors in each particular breed. They are created by the following combinations of the dun gene acting upon an underlying base coat color.
Other variations result from the interplay of additional genes:
A single copy of the cream gene on a black base coat does not significantly lighten black hair, though it may have a subtle effect, and thus a single copy generally has no visible effect on a grullo, either. Conversely, double copies of the cream gene create very light-colored horses (cremello, perlino, and smoky cream). Thus, if a horse with two cream dilution alleles also carries the dun gene, it also will be cream-colored, with primitive markings not visible to any significant degree.
The Fjord horse breed, which is predominantly dun, uses unique Norwegian-based terminology to distinguish between the different shades of dun horses. "Brown dun," or brunnblakk is a zebra dun, rødblakk is a red dun, grå - literally "gray" - is a grullo, buckskin duns are called ulsblakk or white dun, and a dunalino is called a "yellow dun" or gulblakk. A cremello, perlino or smoky cream is called "white" or kvit.
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