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|From Yosemite National Park|
19 spp., see text and map
|From Yosemite National Park|
19 spp., see text and map
The coyote (US // or //, UK //, or //; Canis latrans), also known as the American jackal, brush wolf, or the prairie wolf, is a species of canine found throughout North and Central America, ranging from Panama in the south, north through Mexico, the United States, and Canada. It occurs as far north as Alaska and all but the northernmost portions of Canada. The term is also used for the eastern coyote (Canis latrans var.), which contains not only C. latrans but also significant percentages of Canis lupus lycaon ancestry.
Currently, 19 subspecies are recognized, with 16 in Canada, Mexico, and the United States, and three in Central America. While its closest living relative, the gray wolf, is of Eurasian origin, the coyote has evolved in North America during the Pleistocene epoch 1.8 million years ago (mya), alongside the dire wolf. The coyote fills roughly the same ecological niche in the Americas that is filled in Eurasia and Africa by the similarly sized canids called jackals, which however are, within Canis, neither very closely related to each other, nor the coyote. Unlike the wolf, the coyote's range has expanded in the wake of human civilization, and coyotes readily reproduce in metropolitan areas.
The name "coyote" is borrowed from Mexican Spanish coyote, ultimately derived from the Aztec word cóyotl, meaning "trickster". Its scientific name, Canis latrans, means "barking dog" in Latin.
The color of the coyote's pelt varies from grayish-brown to yellowish-gray on the upper parts, while the throat and belly tend to have a buff or white color. The forelegs, sides of the head, muzzle and paws are reddish-brown. The back has tawny-colored underfur and long, black-tipped guard hairs that form a black dorsal stripe and a dark cross on the shoulder area. The black-tipped tail has a scent gland located on its dorsal base. Coyotes shed once a year, beginning in May with light hair loss, ending in July after heavy shedding. The ears are proportionately large in relation to the head, while the feet are relatively small in relation to the rest of the body. Certain experts have noted the shape of a domestic dog's brain case is closer to the coyote's in shape than that of a wolf's. Mountain-dwelling coyotes tend to be dark-furred, while desert coyotes tend to be more light brown in color.
Coyotes typically grow to 30–34 in (76–86 cm) in length, not counting a tail of 12–16 in (30–41 cm), stand about 23–26 in (58–66 cm) at the shoulder and weigh from 15–46 lb (6.8–20.9 kg). Northern coyotes are typically larger than southern subspecies, with the largest coyotes on record weighing 74.75 pounds (33.91 kg) and measuring 1.75 m (5.7 ft) in total length.
The coyote's dental formula is I 3/3, C 1/1, Pm 4/4, M usually 2/2, occasionally 3/3, 3/2, or 2/3 × 2 = 40, 44, or 42 Normal spacing between the upper canine teeth is 29–35 mm (1.1–1.4 in) and 25–32 mm (0.98–1.26 in) between the lower canine teeth.
The upper frequency limit of hearing for coyotes is 80 kHz, compared to the 60 kHz of domestic dogs. Comparable to wolves, and similar to domestic dogs, coyotes have a higher density of sweat glands on their paw pads. This trait, however, is absent in the large New England coyotes, which are thought to have some wolf ancestry.
Though coyotes have been observed to travel in large groups, they primarily hunt in pairs. Typical packs consist of six closely related adults, yearlings and young. Coyote packs are generally smaller than wolf packs, and associations between individuals are less stable, thus making their social behavior more in line with that of the dingo. In theory, this is due to an earlier expression of aggression, and the fact that coyotes reach their full growth in their first year, unlike wolves, which reach it in their second. Common names of coyote groups are a band, a pack, or a rout. Coyotes are primarily nocturnal, but can often be seen during daylight hours. They were once essentially diurnal, but have adapted to more nocturnal behavior with pressure from humans.
Coyotes are capable of digging their own burrows, though they often prefer the burrows of groundhogs or American badgers. Their territorial ranges can be as much as 19 km in diameter around the den, and travel occurs along fixed trails. Like other canids, coyotes mark their territories with urine.
In areas where wolves have been exterminated, coyotes usually flourish. For example, as New England became increasingly settled and the resident wolves were eliminated, the coyote population increased, filling the empty ecological niche. Coyotes appear better able than wolves to live among people.
Female coyotes are monoestrous, and remain in estrus for two to five days between late January and early March, during which mating occurs. Once the female chooses a partner, the mated pair may remain temporarily monogamous for a number of years. Coyotes also practice alloparental care, in which a coyote pair adopts the pup or pups of another pair. This might take place if the original parents die or are for some reason separated from them. This behavior is common and is seen in many other animal species. Depending on geographic location, spermatogenesis in males takes around 54 days, and occurs between January and February. The gestation period lasts from 60 to 63 days. Litter size ranges from one to 19 pups; the average is six. These large litters act as compensatory measures against the high juvenile mortality rate – about 50–70% of pups do not survive to adulthood. The pups weigh approximately 250 grams at birth, and are initially blind and limp-eared. Coyote growth rate is faster than that of wolves, being similar in length to that of the dhole. The eyes open and ears become erect after 10 days. Around 21–28 days after birth, the young begin to emerge from the den, and by 35 days, they are fully weaned. Both parents feed the weaned pups with regurgitated food. Male pups will disperse from their dens between months 6 and 9, while females usually remain with the parents and form the basis of the pack. The pups attain full growth between 9 and 12 months old. Sexual maturity is reached by 12 months. Unlike wolves, mother coyotes will tolerate other lactating females in their pack.
Coyotes will sometimes mate with domestic dogs, usually in areas such as Texas and Oklahoma, where the coyotes are plentiful and the breeding season is extended because of the warm weather. The resulting hybrids, called coydogs, maintain the coyote's predatory nature, along with the dog's lack of timidity toward humans, making them a more serious threat to livestock than pure-blooded animals. This crossbreeding has the added effect of confusing the breeding cycle. Coyotes usually breed only once a year, while coydogs will breed year-round, producing many more pups than a wild coyote. Differences in the ears and tail generally can be used to distinguish coydogs from domestic or feral dogs or pure coyotes. Breeding experiments in Germany with poodles, coyotes, and later on with the resulting dog-coyote hybrids showed that, unlike wolfdogs, coydogs exhibit a decrease in fertility, significant communication problems, and an increase of genetic diseases after three generations of interbreeding.
Coyotes have also been known, on occasion, to mate with wolves, mostly with eastern subspecies of the grey wolf such as the Great Plains Wolf, though this is less common than with dogs, due to the wolf's hostility to the coyote. The offspring, known as a coywolf, is generally intermediate in size to both parents, being larger than a pure coyote, but smaller than a pure wolf. A study showed that of 100 "coyotes" collected in Maine, 22 had half or more grey wolf ancestry, and one was 89% grey wolf. The large eastern coyotes in Canada are proposed to be actually hybrids of the smaller western coyotes and grey wolves that met and mated decades ago, as the coyotes moved toward New England from their earlier western ranges. Ontario Ministry of Natural Resources research scientist Brent Patterson has revealed findings that most coyotes in Eastern Ontario are wolf-coyote hybrids and that the Eastern wolves in Algonquin Park are, in general, not interbreeding with coyotes.
Similarly, on a population level, Roland Kays, curator of mammals at the State Museum of New York has obtained preliminary DNA evidence for eastern coyotes suggesting interbreeding and a genetic makeup of 85 to 90% coyote, perhaps 10% wolf and slightly less than 5% dog—"a giant Canis soupus", in his words.
The red wolf is thought by some researchers to be in fact a wolf/coyote hybrid rather than a unique species. Strong evidence for hybridization was found through genetic testing, which showed red wolves have only 5% of their alleles unique from either gray wolves or coyotes. Genetic distance calculations have indicated red wolves are intermediate between coyotes and gray wolves, and they bear great similarity to wolf/coyote hybrids in southern Quebec and Minnesota. Analyses of mitochondrial DNA showed the existing red wolf populations are predominantly coyote in origin.
Coyote-Wolf hybrids continue to expand in numbers, and are now being found from Canada all the way down to Virginia.
In an evolutionary biology research conducted by a team of researchers in the Uppsala University, analysis of control region haplotypes of the mitochondrial DNA and sex chromosomes from Mexican grey wolves, a critically endangered subspecies of the grey wolf once nearly driven to extinction in the wild, confirmed the presence of coyote markers in some of the wolves. The study suggests that at some point in time, female coyotes in the south managed to mate with some of the male wolves of the remnant wild Mexican grey wolf populations with the female hybrids backcrossing with other male wolves. Analysis on the haplotype of some coyotes from Texas also detected the presence of male wolf introgression such as Y chromosomes from the grey wolves in the southern coyotes. In one cryptology investigation on a corpse of what was initially labelled as a chupacabra, examinations conducted by the UC Davis team and the Texas State University concluded based on analysis of the mitochondrial and the sex chromosomes that the male animal was in fact another coyote and wolf hybrid sired by a male Mexican wolf.
The calls a coyote makes are high-pitched and variously described as howls, yips, yelps, and barks. These calls may be a long rising and falling note (a howl) or a series of short notes (yips). These calls are most often heard at dusk or night, but may sometimes be heard in the day, even in the middle of the day. Although these calls are made throughout the year, they are most common during the spring mating season and in the fall when the pups leave their families to establish new territories. When a coyote calls its pack together, it howls at one high note. When the pack is together, it howls higher and higher, and then it will yip and yelp and also do a yi-yi sound, very shrill, with the howl.
Sometimes labelled as carnivores but more often as omnivores, coyotes are opportunistic, versatile feeders. They eat small mammals such as (depending on the region in which it lives) voles, prairie dogs, eastern cottontails, ground squirrels, mice, birds, snakes, lizards, deer, javelina, and livestock, as well as insects and other invertebrates. The coyote will also target any species of bird that nests on the ground. They will eat carrion, but tend to prefer fresh meat. Fruits and vegetables can form a significant part of the coyote's diet in the summer and autumn. Part of the coyote's success as a species is its dietary adaptability. As such, coyotes have been known to eat human rubbish and domestic pets. Urban populations of coyotes have been known to actively hunt cats, and to leap shorter fences to take small dogs. In particularly bold urban packs, coyotes have also been reported to shadow human joggers or larger dogs, and even to take small dogs while the dog is still on a leash. This behavior is often reported when normal urban prey, such as brown rats, black rats and rabbits, has become scarce. Yet, confirmed reports of coyotes killing a human have been documented. A 2011 trail camera video uncovered two or three coyotes killing a large deer.
Though the coyote is the basis for the character of Wile E. Coyote in the Warner Brothers Looney Tunes and Merrie Melodies animated cartoons, especially about the Road Runner, coyotes have not been known as yet to successfully attack greater roadrunners for prey.
Coyotes shift their hunting techniques in accordance with their prey. When hunting small animals such as mice, they slowly stalk through the grass, and use their acute sense of smell to track down the prey. When the prey is located, the coyotes stiffen and pounce on the prey in a cat-like manner. They will commonly work in teams when hunting large ungulates such as deer, which is more common in winter (when large prey is likely weakened) and in larger-bodied northern coyotes. Coyotes may take turns in baiting and pursuing the deer to exhaustion, or they may drive it towards a hidden member of the pack. When attacking large prey, coyotes attack from the rear and the flanks of their prey. Occasionally, they also grab the neck and head, pulling the animal down to the ground. Coyotes are persistent hunters, with successful attacks sometimes lasting as long as 21 hours; even unsuccessful ones can continue more than eight hours before the coyotes give up. Depth of snow can affect the likelihood of a successful kill. Packs of coyotes can bring down prey as large as adult elk, which often weigh over 250 kg (550 lbs) or more than 15 times the weight of a fairly large coyote.
The average distance covered in a night's hunting is 2.5 mi (4.0 km).
The gray wolf is a significant predator of coyotes wherever their ranges overlap. Since the Yellowstone gray wolf reintroduction in 1995 and 1996, the local coyote population went through a dramatic restructuring. Until the wolves returned, Yellowstone National Park had one of the densest and most stable coyote populations in America due to a lack of human impacts. Two years after the wolf reintroductions, the population of coyotes had been reduced 50% through both competitive exclusion and intraguild predation compared to before their reintroduction. In Grand Teton, coyote densities were 33% lower than normal in the areas where they coexisted with wolves, and 39% lower in the areas of Yellowstone where wolves were reintroduced. In one study, about 16% of radio-collared coyotes were preyed upon by wolves. Yellowstone coyotes have had to shift their territories as a result, moving from open meadows to steep terrain. Carcasses in the open no longer attract coyotes; when a coyote is chased on flat terrain, it is often killed. They feel more secure on steep terrain, where they will often lead a pursuing wolf downhill. As the wolf comes after it, the coyote will turn around and run uphill. Wolves, being heavier, cannot stop and the coyote gains a large lead. Though physical confrontations between the two species are usually dominated by the larger wolves, coyotes have been known to attack wolves if they outnumber them. Both species will kill each other's pups, given the opportunity. Wolf urine has been marketed and claimed to be an organic coyote deterrent, such as for deterring attacks on sheep.
Cougars sometimes kill coyotes. The coyote's instinctive fear of cougars has led to the development of sound systems which repel coyotes from public places by replicating the sounds of a cougar. Rarely, bears can also kill coyotes, more likely in competitive rather than predatory attacks. However, both cougars and bears have been displaced from carcasses by coyote packs.
In sympatric populations of coyotes and red foxes, fox territories tend to be located largely outside of coyote territories. The principal cause of this separation is believed to be active avoidance of coyotes by the foxes. Interactions between the two species vary in nature, ranging from active antagonism to indifference. The majority of aggressive encounters are initiated by coyotes, and there are few reports of red foxes acting aggressively toward coyotes except when attacked or when their pups were approached. Conversely, foxes and coyotes have sometimes been seen feeding together. In southern California, coyotes frequently kill gray foxes, and these smaller canids tend to avoid areas with high coyote densities.
Coyotes will sometimes form a symbiotic relationship with American badgers. Because coyotes are not very effective at digging rodents out of their burrows, they will chase the animals while they are above ground. Badgers, on the other hand, are not fast runners, but are well-adapted to digging. When hunting together, they effectively leave little escape for prey in the area.
In some areas, coyotes share their ranges with bobcats. It is rare for these two similarly sized species to physically confront one another, though bobcat populations tend to diminish in areas with high coyote densities. However, several studies have demonstrated interference competition between coyotes and bobcats, and in all cases coyotes dominated the interaction. Multiple researchers all reported instances of coyotes killing bobcats, whereas bobcats killing coyotes is more rare. Coyotes attack bobcats using a bite-and-shake method similar to that used on medium-sized prey. Coyotes (both single individuals and groups) have been known to occasionally kill bobcats – in most cases, the bobcats were relatively small specimens, such as adult females and juveniles. However, coyote attacks (by an unknown number of coyotes) on adult male bobcats have occurred. In California, coyote and bobcat populations are not negatively correlated across different habitat types, but predation by coyotes is an important source of mortality in bobcats.
Despite being extensively hunted, the coyote is one of the few medium- to large-sized animals that has enlarged its range since human encroachment began. It originally ranged primarily in the western half of North America, but it has adapted readily to the changes caused by human presence and, since the early 19th century, has been steadily and dramatically extending its range. Sightings now commonly occur in a majority of the United States and Canada. Coyotes inhabit nearly every contiguous U.S. state and Alaska. They have moved into most of the areas of North America formerly occupied by wolves, and are often observed foraging in suburban garbage bins.
Coyotes are difficult to tame, except when raised from a very young age, and even then, much of their wild temperament shows when they reach puberty. Coyotes have never been domesticated with the possible exception of the Hare Indian dogs, which may have been domesticated coyotes or dog-coyote hybrids, used by the Hare Native American tribe of northern Canada for hunting. Naturalist John Richardson, who studied the breed in the 1820s, before it was diluted by crossings with other breeds, could detect no decided difference in form between the breed and coyotes, and surmised it was a domesticated version of the wild animal.
Coyote attacks on humans are uncommon and rarely cause serious injuries, due to the relatively small size of the coyote, but have been increasingly frequent, especially in the state of California. In the 30 years leading up to March 2006, at least 160 attacks occurred in the United States, mostly in the Los Angeles County area. Data from USDA Wildlife Services, the California Department of Fish and Game, and other sources show that while 41 attacks occurred during the period of 1988–1997, 48 attacks were verified from 1998 through 2003. The majority of these incidents occurred in Southern California near the suburban-wildland interface.
In the absence of the harassment of coyotes practiced by rural people, urban coyotes are losing their fear of humans, which is further worsened by people intentionally or unintentionally feeding coyotes. In such situations, some coyotes have begun to act aggressively toward humans, chasing joggers and bicyclists, confronting people walking their dogs, and stalking small children. Non rabid coyotes in these areas will sometimes target small children, mostly under the age of 10, though some adults have been bitten.
Although media reports of such attacks generally identify the animals in question as simply "coyotes," research into the genetics of the eastern coyote indicates those involved in attacks in northeast North America, including Pennsylvania, New York, New England, and eastern Canada, may have actually been coywolves, hybrids of Canis latrans and Canis lupus, not fully coyotes.
Coyotes are presently the most abundant livestock predators in western North America, causing the majority of sheep, goat and cattle losses. For example, according to the National Agricultural Statistics Service, coyotes were responsible for 60.5% of the 224,000 sheep deaths attributed to predation in 2004. The total number of sheep deaths in 2004 comprised 2.22% of the total sheep and lamb population in the United States. According to the National Agricultural Statistics Service USDA report, "All sheep and lamb inventory in the United States on July 1, 2005, totaled 7.80 million head, 2% above July 1, 2004. Breeding sheep inventory at 4.66 million head on July 1, 2005 was 2% above July 1, 2004." Because coyote populations are typically many times greater and more widely distributed than those of wolves, coyotes cause more overall predation losses. However, an Idaho census taken in 2005 showed that individual coyotes were one-twentieth as likely to attack livestock than individual wolves.
Livestock guardian dogs are commonly used to aggressively repel predators and have worked well in both fenced pasture and range operations. A 1986 survey of sheep producers in the USA found that 82% reported the use of dogs represented an economic asset.
Coyotes will typically bite the throat just behind the jaw and below the ear when attacking adult sheep or goats, with death commonly resulting from suffocation. Blood loss is usually a secondary cause of death. Calves and heavily fleeced sheep are killed by attacking the flanks or hindquarters, causing shock and blood loss. When attacking smaller prey, such as young lambs, the kill is made by biting the skull and spinal regions, causing massive tissue and bone damage. Small or young prey may be completely carried off, leaving only blood as evidence of a kill. Coyotes will usually leave the hide and most of the skeleton of larger animals relatively intact, unless food is scarce, in which case they may leave only the largest bones. Scattered bits of wool, skin and other parts are characteristic where coyotes feed extensively on larger carcasses.
Coyote predation can usually be distinguished from dog or coydog predation by the fact that coyotes partially consume their victims. Tracks are also an important factor in distinguishing coyote from dog predation. Coyote tracks tend to be more oval-shaped and compact than those of domestic dogs, and their claw marks are less prominent and the tracks tend to follow a straight line more closely than those of dogs. With the exception of sighthounds, most dogs of similar weight to coyotes have a slightly shorter stride. Coyote kills can be distinguished from wolf kills by the fact that there is less damage to the underlying tissues. Also, coyote scats tend to be smaller than wolf scats.
Coyotes are often attracted to dog food and animals that are small enough to appear as prey. Items such as garbage, pet food, and sometimes feeding stations for birds and squirrels will attract coyotes into backyards. About three to five pets attacked by coyotes are brought into the Animal Urgent Care hospital of south Orange County (California) each week, the majority of which are dogs, since cats typically do not survive the attacks. Scat analysis collected near Claremont, California revealed that coyotes relied heavily on pets as a food source in winter and spring. At one location in Southern California, coyotes began relying on a colony of feral cats as a food source. Over time, the coyotes killed most of the cats, and then continued to eat the cat food placed daily at the colony site by people who were maintaining the cat colony. Coyotes usually attack smaller-sized dogs, but they have been known to attack even large, powerful breeds such as the Rottweiler in exceptional cases. Dogs larger than coyotes are generally able to drive them off, and have been known to kill coyotes. Smaller breeds are more likely to suffer injury or death.
In the early days of European settlement in North Dakota, American beavers were the most valued and sought after furbearers, though other species were also taken, including coyotes. Coyotes are an important furbearer in the region. During the 1983–86 seasons, North Dakota buyers purchased an average of 7,913 pelts annually, for an average annual combined return to takers of $255,458. In 1986–87, South Dakota buyers purchased 8,149 pelts for a total of $349,674 to takers.
The harvest of coyote pelts in Texas has varied over the past few decades, but has generally followed a downward trend. A study from the Texas Parks and Wildlife Department, however, found there was no indication of population decline, and suggested, as pelt prices were not increasing, the decrease in harvest was likely due to decreasing demand, and not increasing scarcity (where pelt prices would go up). It suggested fashion, and the changing custom of wearing fur garments, may be significant among these factors.
Today, coyote fur is still used for full coats and trim and is particularly popular for men’s coats.
Traditional stories from many Native American, First Nations, and Aboriginal cultures include a deity whose name is translated into English as "Coyote". Although especially common in stories told by southwestern Native American nations, such as the Diné and Apache, stories about Coyote appear in dozens of Native American nations from Canada to Mexico.
Usually appearing as a trickster, a culture hero or both, Coyote also often appears in creation myths and etiological myths. Although often appearing in stories as male, Coyote can be female, hermaphrodite, or gender changing, in traditional Aboriginal stories.
The coyote is a popular figure in folklore and popular culture. References may invoke either the animal or the mythological figure. Traits commonly described in pop culture appearances include inventiveness, mischievousness, and evasiveness. By far, the best known representation is the animated Wile E. Coyote, Super Genius, from the Road Runner cartoons, whose popularity has spread the three-syllable Spanish pronunciation of the word coyote throughout English-speaking North America.
The NBA San Antonio Spurs mascot is "The Coyote", as well.
The 19 recognized subspecies of coyote are:
In 1816, in the third volume of Lorenz Oken's Lehrbuch der Naturgeschichte, the author found sufficient similarities in the dentition of coyotes and jackals to place these species into a new separate genus from Canis called Thos after the classical Greek word θώς (jackal). Oken's idiosyncratic nomenclatorial ways, however, aroused the scorn of a number of zoological systematists. Nearly all the descriptive words used to justify the genus division were relative terms without a reference measure, and the argument did not take into account the size differences between the species, which can be considerable. Angel Cabrera, in his 1932 monograph on the mammals of Morocco, briefly touched upon the question of whether or not the presence of a cingulum on the upper molars of the jackals and its corresponding absence in the rest of Canis could justify a subdivision of the genus Canis. In practice, he chose the undivided-genus alternative and referred to the jackals as Canis. A few authors, however, Ernest Thompson Seton being among them, accepted Oken's nomenclature, and went as far as referring to the coyote as American jackal.
The Oken/Heller proposal of the new genus Thos did not affect the classification of the coyote. Gerrit S. Miller still had in his 1924 edition of List of North American Recent Mammals in the section “Genus Canis Linnaeas,” the subordinate heading “Subgenus Thos Oken” and backed it up with a reference to Heller. In the reworked version of the book in 1955, Philip Hershkovitz and Hartley Jackson led him to drop Thos both as an available scientific term and as a viable subgenus of Canis. In his definitive study of the taxonomy of the coyote, Jackson had, in response to Miller, queried whether Heller had seriously looked at specimens of coyotes prior to his 1914 article, and thought the characters to be “not sufficiently important or stable to warrant subgeneric recognition for the group”.
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