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The chelicerae // are mouthparts of the Chelicerata, an arthropod subphylum that includes arachnids, Merostomata (horseshoe crabs), and Pycnogonida (sea spiders). Additionally, some chelicerae, such as those found in spiders, are hollow and contain (or are connected to) venom glands, and are used to inject venom into prey or a (perceived) threat.
Chelicerae can be divided into three kinds: jackknife chelicerae, scissor chelicerae, and 3-segmented chelate chelicerae.
The jackknife chelicera is subchelate (with fixed finger much reduced or absent) and is composed of two segments. This type of chelicera occurs exclusively in the Tetrapulmonata.
Jackknife chelicera presents two different forms: orthognathous and labidognathous. Orthognathous chelicerae are articulated in a manner that enables movements of the appendages parallel to the body axis. This kind of chelicera occurs in the Liphistiomorphae and Mygalomorphae spiders and in the related orders Amblypygi, Schizomida, Thelyphonida. Labidognathous chelicerae moves at right angles to the body axis. This kind of chelicera is rotated and occurs exclusively in the Araneomorphae spiders.
The chelicerae consist of a base segment that articulates with the cephalothorax and a fang portion that articulates with the base segment. Except for the family Uloboridae, the cribellate Orb Weavers, all spiders have venom glands and can inject the venom through openings in the tips of their fangs when biting prey. The glands that produce this venom are located in the two segments of the chelicerae, and, in most spiders, extend beyond the chelicerae and into the cephalothorax.
The fang, the organic functional equivalent to a hypodermic needle is what penetrates the skin, fur, or exoskeleton of the spider's target—spider mouthparts are primarily intended for envenoming a spider's prey in most species, typically insects and other small arthropods. The basal portion includes all or part of the spider's venom glands, which can be squeezed to control the amount of venom forced out of the glands. Such control permits a spider to administer either a dry bite, a dose appropriate to the nature of the prey or enemy, or a maximal dose. The control is also necessary for actions such as the spitting of venomous silk by members of the family Scytodidae; they depend on that mechanism both in hunting and defence.
When a spider bites, the two parts of the chelicerae come together like a folding knife, and when making a threat display or actually preparing to bite, the spider will open the angle of the fangs with the basal portion of the chelicerae and also open the angle of the basal portion with the cephalothorax. In the tarantulas and other Mygalomorphae, the horizontal separation of the tips of the fangs does not change much, but in the other spiders the tips of the fangs move apart from each other as well as elevating. Even the tips of the fangs of the rather large spider shown above are quite sharp, and the spider's body is well adapted to driving the fangs into flesh. Some spider bites, such as those of the Sydney funnel-web spider, are reported to have penetrated toenails and soft leather shoes.
This is the primitive condition and occurs in arachnids such as the Scorpiones and the Opiliones, as well as in non-arachnid Chelicerata such as the Xiphosura and Eurypterida. The chelifores of the Pycnogonida may be homologous to chelicerae.